Evergreen shrub to moderate-sized tree to 6(-10) m tall, with brownish to gray, longitudinally furrowed bark; twigs slightly to moderately angled, slender, sparsely to moderately lepidote, otherwise subglabrous to slightly (moderately) pubescent; buds ovoid, 0.8-2.5 x 0.8-2 mm. Leaf blades elliptic to ovate, obovate, or nearly orbicular, 1.7-6.5(-9) x 0.7-4(-5.2) cm, ± flat to slightly abaxially curved, coriaceous, ca. 0.2-0.36 mm thick; base narrowly cuneate to rounded (very slightly cordate); apex acuminate, to acute, rounded, or truncate; margin plane to slightly revolute, the apical portion strongly to obscurely and irregularly toothed to entire, the basal portion entire to irregularly and obscurely toothed; venation brochidodromous, 3° veins ± reticulate; adaxial surface lepidote but scales usually quickly deciduous, sparsely to moderately pubescent on midvein, especially basal portion, the 3° and higher order veins visible to obscure, the 2° veins prominent to obscure, sometimes depressed; abaxial surface moderately lepidote, otherwise glabrous (sparsely pubescent on midvein only, near petiole), the 3° and higher order veins obscure to very slightly raised, not forming dense, fine, raised network, the 2° obscure to slightly raised and visible; scales rust colored, persistent to deciduous, ca. 0.1-0.34 mm in diameter, erose to entire; petiole 2-10 mm long, lepidote, otherwise pubescent adaxially or all around; flower buds ± intermixed with vegetative buds. Inflorescences fasciculate, 4- to 16-flowered; pedicels clearly articulated with calyx, slender, 4-11 mm long, lepidote, otherwise glabrous to densely pubescent; bracts ± opposite, basal or nearly so, narrowly triangular, 0.9-2 mm long; floral bracts to ca. 2 mm long. Flowers 4- or 5-merous; calyx lobes triangular, with acuminate apices, 0.7-1.5(-2) x 0.6-1.3 mm, adaxial side glabrous or sparsely pubescent, especially near apex, abaxial side lepidote, otherwise glabrous to sparsely pubescent; corolla cylindrical, white (pink tinged toward mouth), 4-8.5 x (2-)2.5-4 mm, abaxially sparsely lepidote; filaments roughened, 3-5 mm long, unappendaged; anthers 0.7-1.5 mm long; ovary lepidote, otherwise pubescent, placentae ± subapical. Capsules ellipsoid to ovoid, (3-)3.5-6 x 1.7-3.5(-4) mm, slightly lepidote, otherwise sparsely pubescent, especially near base, the pale, very thick sutures separating as unit from adjacent valves; seeds 1.5-3 mm long.
Distribution. Hispaniola and Puerto Rico, in
open forests of Pinus occidentalis, thickets, and cloud forests
of various mountain ranges, at ca. (500-)1100-2100(-2300) m elev.
Key to the Varieties of Lyonia stahlii
1. Leaves entire to obscurely (occasionally clearly) toothed,
secondary
veins obscure to slightly depressed; flowers
mainly 5-merous,
corolla 4-6 mm long; Puerto Rico ..............................
var. stahlii.
1. Leaves irregularly toothed or notched, secondary veins often
depressed; flowers 5- and/or 4-merous,
corolla (5.5-)6-8.5 mm
long; Hispaniola ........................................................
var. costata.
Lyonia stahlii Urban var. stahlii, Symb. antill. 5: 453. 1908. Xolisma stahlii (Urban) Small, N. Amer. Fl. 29(1): 71. 1914. Lyonia rubiginosa (Persoon) G. Don var. stahlii (Urban) Judd, J. Arnold Arbor. 62: 355. 1981. Lyonia rubiginosa (Persoon) G. Don subsp. stahlii (Urban) Borhidi, Acta Bot. Hung. 29: 188. 1983. Illustrtion: Judd (1995), fig. 14. Type. Puerto Rico, nr. Bayamón, in Monte Helechol, 500 m, 1885, A. Ståhl 468 (holotype, B, n.v., destroyed; frag. of holotype, NY). Image: Habit.
Shrub to moderate-sized and often densely branched tree to 6(-10) m tall; twigs subglabrous to slightly (moderately) pubescent. Leaf blades 1.7-6.5(-9) x 0.8-4(-5.2) cm, margin apically obscurely and irregularly toothed to entire (strongly toothed), basally entire to obscurely toothed; adaxial surface with the 3° and higher order veins faintly visible to obscure, the 2° veins obscure to visible, usually not depressed or only slightly so. Pedicels slender, 4-11 mm long. Flowers (4- or)5-merous; corolla 4-6 x (2-)2.5-4 mm. Capsules 3.5-5.5 x 2-3.5(-4) mm.
Distribution. Endemic to Puerto Rico (Cordillera
Central), in moist cloud forests, at ca. (500-)1100-1338 m elev.
Locally common on highest peaks of range (Cerro de Punta, Monte Jayuya,
Cerro Maravillas, Pico Guilarte) in dense thickets or cloud forests.
Also in disturbed roadside vegetation; seedlings common on road-cuts or
landslides (if near seed-source). Flowering January through August.
Lyonia stahlii Urban var. costata (Urban) Judd, Fl. Neotrop. Monogr. 66: 263. 1995. Lyonia costata Urban, Symb. antill. 7: 316. 1912. Xolisma costata (Urban) Small, N. Amer. Fl. 29(1): 71. 1914. Lyonia rubiginosa (Persoon) G. Don var. costata (Urban) Judd, J. Arnold Arbor. 62: 354. 1981. Lyonia rubiginosa (Persoon) G. Don subsp. costata (Urban) Borhidi, Acta Bot. Hung. 29: 188. 1983. Illustration: Judd (1995), fig. 13. Type. Dominican Republic. Between Constanza and Valle Nuevo, 1700 m, Apr 1910, von Tuerckheim 3119 (holotype, B, n.v., destroyed; isotypes, F, GH, M, MO, NY, L, P, S, US). Image: Habit.
Lyonia acutifolia Urban & Ekman in Urban, Ark. Bot. 20A: 35. 1926. Type. Haiti. Massif de la Selle, Morne Tranchant, Pétionville, on the ridge toward Godet, ca. 1850 m, 19 Sep 1924, Ekman H1949 (holotype, S; isotypes, GH, IJ, K, NY, S, US).Erect shrub to moderate-sized tree to 4(-10) m tall. Twigs subglabrous to slightly pubescent. Leaf blades 2-6(-8.5) x 0.7-3(-5) cm; margin apically strongly to obscurely and irregularly toothed, basally entire to obscurely toothed; adaxial surface with 3° and higher order veins faintly visible to obscure, usually at least some 2° veins visible and slightly to moderately depressed. Pedicels slender, 5-10 mm long. Flowers 4- or 5-merous; corolla (5.5-)6-8.5 x (2-)2.5-4 mm. Capsules (3-)3.5-6 x 1.7-3.5(-4) mm.Lyonia angulata Urban & Ekman in Urban, Ark. Bot. 20A: 35. 1926. Type. Haiti. Massif de la Selle, Morne Cabaio, Jardin Bois Pin, ca. 1900 m, 25 Aug 1924, Ekman H1632 (holotype, S; isotypes, IJ, S, US).
Lyonia darrasiana Urban & Ekman in Urban, Ark. Bot. 21A: 52. 1927. Type. Haiti. Massif du Nord, nr. La Borgne, top of Morne Darras, ca. 1175 m, 3 Sep 1925, Ekman H4771 (holotype, S; isotypes, IJ, S).
Xolisma longelaminata Sleumer, Repert. Sp. Nov. 36: 271. 1934. Lyonia longelaminata (Sleumer) Jiménez, Rhodora 62: 237. 1960. Type. Dominican Republic. Santiago: Cordillera Central, Monción, Monte Gallo, ca. 1775 m, 10 Jun 1929, Ekman H12917 (holotype, S; isotypes, A, GH, NY).
Distribution. Endemic to Hispaniola (Cordillera
Central, Massif du Nord, Sierra de Neiba, Massif de la Selle, and Sierra
de Baoruco, including Monteada Nuevo), in low-elevation Pinus occidentalis
forests, thickets, and cloud forests, at (900-)1100-2100(-2300) m elev.
Flowering probably late March through early May.